No idea... but the small HD version of Trailer 2 opened the Quicktime application for me and started buffering content; it looked like it was going to play, though I wasn't patient enough to wait for it to finish. (OSX 10.5.8, QT 7.6.4, Firefox 3.5.5.) (Trailer 1 just opened a pop-up window and started buffering content; it didn't open the Quicktime application. Trailer 2 non-HD did the pop-up with question mark that I'm assuming you saw.)
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Very odd.